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番号
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学名 / 和名
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分布
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**
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*
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備考
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Tribe Ateuchini Perty, 1830
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|
1
|
Paraphytus dentifrons Lewis, 1895 / ダルマコガネ
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T-Nak, Amm, Kkr, Tok, Okn
|
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II-64-354
|
|
|
Tribe Coprini Leach, 1815
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|
2
|
Copris (Copris) acutidens Motschulsky, 1860 / ゴホンダイコクコガネ
|
HOK, Oks, HON, Sad, Awj, Its, SHI, Shd, KYU, Tsm, Got, Krs; Korea, Jeju, China, Taiwan?
|
●
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II-64-355
|
|
|
3-1
|
Copris (Copris) brachypterus brachypterus Nomura, 1964 / マルダイコクコガネ 基亜種
|
Amm
|
|
II-64-355
|
絶滅危惧 II類（VU）
|
|
3-2
|
Copris (Copris) brachypterus toguchii Ochi & Kawahara, 2012 / マルダイコクコガネ 徳之島亜種
|
Tok
|
|
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絶滅危惧 II類（VU）
|
|
4
|
Copris (Copris) ochus (Motschulsky, 1860) / ダイコクコガネ
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HOK, HON, Sad, Iz-O, Iz-Shi, Iz-My, Oki, SHI, KYU, Iki, Tsm, Got, Yak, Kue; Korea, Jeju, China, Mongolia
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●
|
II-64-354
|
絶滅危惧 II類（VU）
|
|
5
|
Copris (Copris) pecuarius Lewis, 1884 / ミヤマダイコクコガネ
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HON, SHI, KYU, Tsm; Korea, Siberia(east)
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●
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II-64-354
|
|
|
6
|
Copris (Copris) tripartitus C. O. Waterhouse, 1875 / ヒメダイコクコガネ
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HON, Tsm; China, Korea, Jeju, Taiwan
|
|
II-64-355
|
情報不足（DD）
|
|
Tribe Deltochilini Lacordaire, 1855
|
|
7
|
Panelus kubotai Kawahara, Inagaki & Ochi, 2007 / ヤクシマアカチャダルマコガネ
|
Yak
|
|
|
|
|
8
|
Panelus ovatus Nomura, 1973 / マルダルマコガネ
|
Kan, KYU, T-Kch, T-Nak, T-Sw, T-Aks, Amm, Tok
|
|
II-354
|
|
|
9
|
Panelus parvulus (Waterhouse, 1874) / マメダルマコガネ
|
HON, Aws, Sad, Iz-My, Iz-Mk, Kan, Awj, Its, Oki, Yas, SHI, Shd, KYU, Tsm, Shk, Cokn, Nkn, Got, Amk, KKs, NKs, SKs, Yak
|
●
|
II-64-354
|
|
|
10
|
Panelus rufulus Nomura, 1973 / アカダルマコガネ
|
Ish, Iri, Hat
|
|
II-64-354
|
|
|
Tribe Oniticellini Kolbe, 1905
|
|
11
|
Liatongus minutus (Motschulsky, 1860) / ツノコガネ
|
HOK, HON, Sad, SHI, KYU, Tsm; Korea, Jeju, China, Taiwan
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●
|
II-64-355
|
Liatongus phanaeoides (Westwood, 1840)
|
|
Tribe Onthophagini Streubel, 1846
|
|
12
|
Caccobius (Caccobius) brevis C. O. Waterhouse, 1875 / ヒメコエンマコガネ
|
HON, Sad, Oki, SHI, Okns, KYU, Iki, Tsm, Got, Yak, Kue; Siberia(east), China(north), Korea, Jeju
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●
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II-65-356
|
Caccobius matsuoi Matsumura & Yohena, 1937
|
|
13
|
Caccobius (Caccobius) jessoensis Harold, 1867 / マエカドコエンマコガネ
|
HOK, Rsr, Oks, HON, Sad, Awj, SHI, KYU
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●
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II-64-356
|
Caccobius hirayamai Matsumura, 1936; Caccobius sapporensis Matsumura, 1936; Caccobius yubaeiensis Matsumura, 1936; Caccobius
amagisanus Matsumura, 1937
|
|
14
|
Caccobius (Caccophilus) nikkoensis (Lewis, 1895) / ニッコウコエンマコガネ
|
HON, SHI, KYU, Tsm
|
●
|
II-65-357
|
Caccobius kasuganus Matsumura & Yohena, 1936
|
|
15
|
Caccobius (Caccophilus) suzukii Matsumura, 1936 / スズキコエンマコガネ
|
HON, KYU; Korea
|
|
II-64-356
|
Caccobius (Caccophilus) boucomonti Balthasar, 1935
|
|
16
|
Caccobius (Caccophilus) unicornis (Fabricius, 1798) / チビコエンマコガネ
|
HON, SHI, KYU, Iki, Okn; Korea, China, Taiwan, Philippines, Malay Pen., Indochina, India, Sri Lanka
|
●
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II-65-357
|
Caccobius yamauchii Matsumura, 1936
|
|
17
|
Digitonthophagus gazella (Fabricius, 1787) / ガゼラエンマコガネ
|
Ihy; Asia, Micronesia, Meranesia, Oceania, Africa, N-America
|
|
|
人為分布種
|
|
18-1
|
Onthophagus (Gibbonthophagus) amamiensis amamiensis Nomura, 1965 / ウシヅノエンマコガネ 基亜種
|
Amm, Tok
|
|
II-65-360
|
|
|
18-2
|
Onthophagus (Gibbonthophagus) amamiensis matsumurai Ochi, Kawahara & Masumoto, 2019 / ウシヅノエンマコガネ 沖縄亜種
|
Okn
|
|
II-65-360
|
|
|
19
|
Onthophagus (Gibbonthophagus) apicetinctus d'Orbigny, 1898 / ヤエヤマコブマルエンマコガネ
|
Ish, Iri, Yon
|
|
II-65-360
|
|
|
20
|
Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 / コブマルエンマコガネ
|
HOK, HON, Tob, Sad, Iz-My, Iz-Mk, Iz-Hch, Kan, Awj, Its, Oki, SHI, Shd, Okns, KYU, Tsm, Shk, Nkn, Got (Wak), KKs, NKs, SKs, Tan, Yak, Mgs, T-Nak, T-Tak, Okn;
Siberia(east), China, Krea, Jeju
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●
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II-65-360
|
Onthophagus akirai Matsumura, 1937; Onthophagus ibonus Matsumura, 1937
|
|
21
|
Onthophagus (Gibbonthophagus) proletarius Harold, 1875 / ヨツモンエンマコガネ
|
Ish; Taiwan, China, Indochina, Malay
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|
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移入種
|
|
22
|
Onthophagus (Gibbonthophagus) viduus Harold, 1874 / マルエンマコガネ
|
HOK, Oks, HON, Tob, Sad, Iz-O, Iz-N, Iz-My, Iz-Hch, Awj, Oki, SHI, KYU, Tsm, Got, Tan, Yak, Kue, T-Nak, T-Kdk, T-Tak, Amm, Tok, Kkr, Kik, Okn, Ihy, Izn, Tnk, Agn, Sng,
Ouj, Tks, Zmm, Kum, Myk, Mnn, Trm, Irb, Smj, Ish, Iri, Yon, Tkt, Khm, Kur, Htm, Hat; Siberia(east), China, Korea, Jeju
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●
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II-65-360
|
Caccobius jononis Matsumura, 1936; Onthophagus aequiperus Matsumura & Yohena, 1937;
Onthophagus chibanuus Matsumura & Yohena, 1937;
Onthophagus hyuganus Matsumura, 1937; Onthophagus komabellus Matsumura, 1937; Onthophagus misujianus Matsumura, 1937
|
|
23
|
Onthophagus (Hikidaeus) shirakii Nakane, 1960 / ネアカエンマコガネ
|
Amm, Kkr, Tok
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|
II-65-358
|
Onthophagus (Onthophagus) shirakii Nakane, 1960
|
|
24
|
Onthophagus (Indachorius) suginoi Ochi, 1984 / ヤンバルエンマコガネ
|
Okn, Kum
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|
II-65-359
|
|
|
25
|
Onthophagus (Matashia) lutosopictus Fairmaire, 1897 / アカマダラエンマコガネ
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Trm, Ish, Tkt, Kur
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|
II-65-360
|
|
|
26
|
Onthophagus (Matashia) ohbayashii Nomura, 1939 / ナガスネエンマコガネ
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HON, Kan, SHI, KYU, Yak; China, Korea, Jeju
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●
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II-65-359
|
|
|
27
|
Onthophagus (Matashia) solivagus Harold, 1886 / キモンエンマコガネ
|
HON; Siberia(east), China, Korea
|
|
II-65-359
|
CPCに日本の分布なし
|
|
28
|
Onthophagus (Onthophagus) bivertex Heyden, 1887 / シナノエンマコガネ
|
HOK, HON, Oki, SHI, KYU, Tsm; Kuril, Siberia(east), Koera, Jeju, China
|
●
|
II-65-357
|
|
|
29
|
Onthophagus (Palaeonthophagus) gibbulus gibbulus (Pallas, 1781) / チャバネエンマコガネ
|
HOK, HON; Siberia, Sakhalin, China(northeast), Korea, Central Asia, Asia Minor, Europe
|
|
II-65-357
|
絶滅危惧 II類（VU）
|
|
30
|
Onthophagus (Palaeonthophagus) ocellatopunctatus C. O. Waterhouse, 1875 / アラメエンマコガネ
|
HOK, HON, Iz-O
|
|
II-65-357
|
準絶滅危惧（NT）
|
|
31
|
Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 / ウエダエンマコガネ
|
HON; Siberia, China(north), Korea
|
|
II-65-358
|
Onthophagus kandai Matsumura, 1937
|
|
32
|
Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 / トビイロエンマコガネ
|
Yon; China, Taiwan, Indochina, Southeast Asia
|
|
II-65-359
|
|
|
33
|
Onthophagus (Paraphanaeomorphus) trituber (Wiedemann, 1823) / ミツコブエンマコガネ
|
HON, Myk, Ogm, Irb; Taiwan, China, Indochina, Indonesia, Malaysia
|
●
|
|
移入種
|
|
34
|
Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 / クロマルエンマコガネ
|
HOK, Oks, HON, Aws, Sad, Iz-My, Iz-Mk, Iz-Hch, Awj, Its, Oki, SHI, Shd, KYU, Tsm, Iki, Got, Yak; Siberia(east), Korea, Jeju, China, Taiwan
|
●
|
II-65-359
|
Onthophagus chuzenjianus Matsumura, 1937; Onthophagus kawarinus Matsumura, 1937
|
|
35
|
Onthophagus (Phanaeomorphus) fodiens Waterhouse, 1875 / フトカドエンマコガネ
|
HON, Sad, Its, Oki, SHI, Shd, KYU, Nkn, Got, SKs, Yak; China, Korea, Jeju
|
●
|
II-65-359
|
Onthophagus hikosanus Matsumura, 1937
|
|
36
|
Onthophagus (Strandius) japonicus Harold, 1874 / ヤマトエンマコガネ
|
HON, Sad, SHI; Siberia(east), China, Korea, Jeju, Taiwan
|
●
|
II-65-358
|
準絶滅危惧（NT）
|
|
37
|
Onthophagus (Strandius) lenzii Harold, 1874 / カドマルエンマコガネ
|
HOK, HON, Aws, Sad, Iz-O, Iz-T, Iz-N, Iz-My, Iz-Mk, Iz-Hch, Iz-Hk, Iz-Ao, Kan, Awj, Its, Oki, SHI, Shd, KYU, Tsm, Iki, Got (Nkd), KKs, SKs, Yak, Mgs, Krs, Kue, T-Nak,
T-Gaj, T-Sw, T-Aks, Kik; China, Korea, Jeju, Taiwan
|
●
|
II-65-358
|
|
|
38
|
Onthophagus (Strandius) oshimanus Nakane, 1960 / オオシマエンマコガネ
|
Amm, Tok
|
|
II-65-358
|
|
|
39
|
Onthophagus (Strandius) yakuinsulanus Nakane, 1984 / ヤクシマエンマコガネ
|
Yak
|
|
|
絶滅危惧 II類（VU）
|
|
40
|
Parascatonomus (Necramator) acuticollis sakishimanus (Nomura, 1976) / トガリエンマコガネ
|
Ish, Iri; Taiwan, China
|
|
II-65-362
|
Onthophagus (Parascatonomus) acuticollis sakishimanus Nomura, 1976
|
|
41
|
Parascatonomus (Necramator) aokii (Nomura, 1976) / ヨナグニエンマコガネ
|
Yon
|
|
II-65-362
|
Onthophagus (Parascatonomus) aokii Nomura, 1976
|
|
42-1
|
Parascatonomus (Necramator) itoi itoi (Nomura, 1976) / オキナワエンマコガネ 基亜種
|
Okn, Ihy
|
|
II-65-362
|
Onthophagus (Parascatonomus) itoi Nomura, 1976
|
|
42-2
|
Parascatonomus (Necramator) itoi kumejimensis Ochi, Kawahara & Masumoto, 2019a / オキナワエンマコガネ 久米島亜種
|
Kum
|
|
|
|
|
43-1
|
Parascatonomus (Necramator) murasakianus murasakianus (Nomura, 1976) / ムラサキエンマコガネ 基亜種
|
Trm, Ish, Iri, Tkt, Khm,
|
|
II-65-361
|
Onthophagus (Parascatonomus) murasakianus murasakianus Nomura, 1976
|
|
43-2
|
Parascatonomus (Necramator) murasakianus carnarius (Nomura, 1976) / ムラサキエンマコガネ 沖縄・奄美亜種
|
T-Nak, T-Aks, T-Tak, Amm, Kkr, Tok, Okn, Ihy, Tnk, Tks, Zmm, Kum,
|
|
II-65-361
|
Onthophagus carnariu Nomura, 1976 / ヒメツヤエンマコガネ;
Onthophagus (Parascatonomus) murasakianus carnarius Nomura, 1976
|
|
43-3
|
Parascatonomus (Necramator) murasakianus miyakoinsularis (Ochi, Y. Miyake & Kusui, 1999) / ムラサキエンマコガネ 宮古島亜種
|
Myk, Irb, Trm,
|
|
|
Onthophagus (Parascatonomus) carnarius miyakoinsularis Ochi, Y.Miyake & Kusui, 1999 / ヒメツヤエンマコガネ
|
|
44
|
Parascatonomus (Necramator) nitidus (Waterhouse, 1875) / ツヤエンマコガネ
|
HOK, HON, Tob, Aws, Sad, Iz-T, Iz-N, Iz-Shi, Iz-Kz, Iz-My, Iz-Mk, Kan, Its, Oki, Yas, SHI, Shd, Okns, KYU, Iki, Shk, Cokn, Nkn, Got (Fuk, Nkd, Wak), NKs, SKs, Tan,
Yak, Mgs, Krs, T-Aks, Amm
|
●
|
II-65-361
|
Onthophagus (Parascatonomus) nitidus Waterhouse, 1875 / ツヤエンマコガネ; Onthophagus (Parascatonomus) nitidus yakushimanus Nomura, 1976
|
|
45
|
Parascatonomus (Necramator) shibatai (Nakane, 1960) / アマミエンマコガネ
|
Amm, Kkr, Tok
|
|
II-65-362
|
Onthophagus (Parascatonomus) shibatai Nakane, 1960
|
|
46
|
Parascatonomus (Necramator) toricornis (Wiedemann, 1823) / ミツノエンマコガネ
|
HON, SHI, KYU, Iki, Nkn; China, Taiwan, Java, Malay Pen., Indochina, Burma, India
|
●
|
II-65-361
|
Onthophagus (Parascatonomus) toricornis (Wiedemann, 1823)
|
|
47
|
Parascatonomus (Parascatonomus) miyakei (Ochi & Araya, 1992) / オオツヤエンマコガネ
|
Ish, Iri; Taiwan, China, Vietnam, Thai
|
|
II-65-361
|
Onthophagus (Parascatonomus) discedens Sharp, 1875; Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992
|
|